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Phylum Archeocyathida
Range: Lower
to Upper Cambrian
Some place this phylum under the sponges - structurally and
functionally similar - others place this group as a separate phylum.
1. organisms with calcareous (microgranular calcite) skeleton
2. invested cup erect or curved, held to substrate with holdfast
3. skeleton may consist of a single porous wall - Monocyathida
more commonly as two concentric porous walls,
an inner and
outer wall separated by a space (intervallum)
4. inner wall surrounds a central cavity
5. intervallum may contain various radial elements:
porous septa - subdivide intervallar space tabulae
- transverse porous skeletal elements of variable curvature
non-porous bars, rods, dissepiments
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Sponges: Phylum
Porifera
Latin porus - pore fera - to bear Range: PreCambrian - Recent
Characteristics: 1. cellular grade organization 2. dominantly marine 3. metazoa 4. asymmetrical or radially symmetrical 5. without organs, mouth or nervous system 6. with a body permeated with pores, canals and chambers (inlets and exits.) 7. water flows through openings by action of numerous flagellate cells that line that internal cavities 8. Larvae free swimming Adults sessile 9. Reproduction usually sexual; freshwater sponges asexual 10. internal skeleton crystalline spicules - Si02 organic fibers
Phylum Porifera (Tasch, 1980)
CLASS 1. Demospongea Range Cambrian to Recent skeletons spongin, siliceous spicules; spicule rays meet at 60o or 120o
CLASS 2. Hyalospongea Range Pre-Cambrian to Recent siliceous spicules - skeletal spicule rays form right angles common spicule stauract - four rays in one plane
CLASS 3. Calcispongea Range Cambrian to Recent calcareous spicules; diacts, triacts, tetracts (tuning fork type spicules may be interlocking to form)
CLASS 4. Sclerospongea Range Ord - Permian (?) Tr - Rec coralline sponge with compound skeleton of siliceous spicules, organic fibers and aragonite
Phylum Porifera: (Boardman, 1987)
Subphylum - Symplasma
CLASS - Demospongia (C - Rec) knobby siliceous spicules fused together. Girtycoelia. Burgess Shale
CLASS - Hexactinellida (C - Rec) six-rayed spicules, more advanced forms have fused spicules to make a three- dimensional rectilinear pattern. Hydnoceras
Subphylum - Cellularia
CLASS - Calcarea (L Perm - Rec) calcium carbonate spicules. Reef formers in the late Paleozoic. Astreosponggium, Stellispongia, Peronidella
CLASS- Sclerospongea (ord-Perm (?), Tr - Rec) coraline sponge with compound skeleton of siliceous spicules, organic fiber and aragonite
CLASS - Stromatoporata* (?C, Ord-K)
CLASS- Chaetetida* (Ord - Rec May belong to Demospongia or Calcarea
CLASS (Order) Sclerospongia
New class; Hartman & Goreau, 1970; symp. Zool. Soc. London, 1969, v. 25, p. 205-243
Group of living sponges (coralline sponges)basal skeleton mass of aragonite (all other calc. sponges calcite), also contain silica and spongin fibersfrom Christmas Islands, Cuba, Jamaica; flourish in deep shade under fore-reef projections (10m - 100m and deeper), twilight to total darkness.
Merlia (similar to certain tabulates of Chaetitidae (sponges?)), Ceratoporella, Stromatospongia, Goreaviellabeige, red vermillion colorsymbiotic with serpulids imbedded and attachedrapid growth
Morphology: elongate depressions converging on osculum, transparentAffinities: stromotoporoids, chaetitids
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