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Phylum SARCODINA
Protozoans: shell given the name TEST
Introduction:
1. simplest & most primitive members of animal kingdom
protos - first zoon - animal
2. range - Cambrian to Recent
3. generally small - 1 micron to 19cm
4. consist of a single cell
cytoplasm; nucleus; cell wall; vacuoles
motion by cilia, flagellum, or pseudopodia
5. water, waste, and 02 distributed by osmosis
6. lack systems, but are completely independent organisms
(organelles)
7. mostly solitary, but a few are colonial in protoplasm matrix
8. feed on other protozoans and microscopic plants, plus organic
debris
some parasitic
9. reproduction asexual or sexual - alternation of generations.
Often produce
spores (resting stages)
10. many have some kind of external protection - hard parts
silica - Radiolarians
chiton (tectin)
agglutinated material
calcium carbonate
11. inhabit fresh, brackish, salt water, moist earth, digestive
& circulatory systems
major food supply of oceans
13. important rock formers locally
Globigerina and radiolarian oozes
fusulinid Ls of Penn. - Perm.
chalks of Cretaceous & Tertiary
Salem Ls of Indiana (Endothyrid forams)
14. very important as index fossils - especially in the petroleum
industry
ORDER FORAMINIFERA
1. possess
netlike pseudopodia (extensions of protoplasm)
2. make up 2 1/2% of all know animals,
fossil or living;
88% of the forams known as fossils
3. mostly benthonic; shallow water to 30,000'
4. some adapted planktonic habit in the Jurassic
(vacuolated protoplasm, spines, reduction of S. G.)
5. density to 2,500 per cm2 on sea floor
Skeletal features Shell termed a TEST
1. Allogromids: Tectin - proteinaceous substance resembling
chiton (C-Hol)
2. Textularids: Arenaceous (agglutinated) - tests of foreign
material. May be highly selective as to type of material
cements calcareous, siliceous, ferruginous (C - HOL)
3. Calcareous -
Rotalina: Hyaline (perforate) - (Permian - Hol)
glassy, hole-bearing (perforate),
calcite and/or
aragonite arranged in
one or more layers
Miliolina: Porcellanous (imperforate) - (Miss
- Hol)
chalky, lack holes, tiny
calcite needles, calcite xls
parallel to outer surface
layer, smooth, shiny, milky
white surface
Fusulinina: Microgranular - (Ord - Tr)
randomly arranged, equidimensional
grains of calcite
sugary appearance.
See P. 82 of Boardman/Cheetham/Rowell
Location and shape of aperture Shape of test
figure 2.3 (Moore. et al.) figure 2.5 (Easton)
figure 2.6 (Easton)
Ornamentation
Perforate
Imperforate
smooth surface - ornamentation
costae - ribs or ridges
granules or nodes
spines - usually planktonic
flanges
pits
reticulate - network of fine lines
Test size
micro - 0.001mm
small - few chambers (0.4mm)
larger - many chambers (2mm and larger)
Dimorphism - 2 body forms in the same species
megalospheric - asexual, few chambers, large proloculus; small test
microspheric - sexual, many chambers, small proloculus; large test
alteration of generations
Key to Superfamilies
I. Test arenaceous (Cambrian to Recent) A. Non-chambered . . . . . . . . . . . . . . . . . . . . . . . . Astrorhizicae
B. Chambered . . . . . . . . . . . . . . . . . . . . . . . . . . . . Lituolicae
II. Test Calcareous A. Mostly imperforate, non hyaline 1. discoid to fusiform, granular . . . . Endothyricae 2. Milioline, porcelanous . . . . . . . . . Miliolicae B. Mostly perforate, hyaline
- chambers tend to lie in linear or planispiral series,
particularly in early portions . . . Lagenicae 2. chambers tend to lie in complex whorls a. tests mostly helicoid around a main long axis . . . . Buliminicae b. tests mostly rotalioid . . . . Rotaliicae
Astrorhizicae Arenaceous tubes which are not divided into chambers. Recovered from insoluble residues from Ls which has been dissolved in dilute acid.
Lituolicae Chambered arenaceous forams Differentiated according to chamber distribution Sil - Hol, very common in K
Endothyricae Catch-all for granular, calcareous, planispiral forms A few arenaceous forms Fusulinids - larger forams, dominated the late Paleozoic Extremely important as index fossils Dev.(?) Miss. - Perm. primitive forms discoidal with rounded chambers, simple walls typified by Endothyra from the Salem Ls, Indiana fusiform group evolved from the Endothyrids during Chester (Miss.) to give fusulinids.
Axial and equatorial sections required for I.D. wall structure of fusulinids
Profusulinella
Fusulina
Schwagerina
Evolutionary trends of the Fusulinids rapid changes in Penn. & Perm. change in shape - discoidal to spherical, spindle-shaped, or subcylindrical increase in size - fraction of an inch in Miss, up to 60mm in Permian complication of shell-well structure - development of thicker primary walls and fewer wall layers fluting of septa development of special structures (chromata, septula, axial fillings) variety of internal structures
1. Resorbed CaC03 tunnels septal pores foramen 2. Redeposited CaC03 chomata - thickened zones adjacent to tunnels axial filling - solid central region - maximum diameter depositional sites for resorbed CaC03 adjustment of S.G. with continued growth
Importance of Fusulinids 1. Restricted geologic range - Miss. to Perm. 2. Abundant & well preserved - fusulinid Ls 3. Widely spread geographically - all continents except Australia & Antarctica 4. Evolution can be traced - thin section 80 million year life span 72 genera, >1000 species
Miliolicae calcareous, (porcelaneous), imperforate most coiled resemble seeds surrounded by husks Triassic to Holocene; abundant after mid-Cretaceous Megalospheric & Microshperic tests; show alternation of generations developmental series starts with: quinqueloculine shape, passes through a trioculine stage, and terminates in a biloculine stage tests resemble long-extinct fusulinids
Lagenicae calcareous, hyaline, perforate basic shape flask-like; also uniserial & spiral Cambrian to Recent, insignificant until Jr., when they were very common through the Cretaceous. Still important.
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