Phylum SARCODINA

Protozoans: shell given the name  TEST

Introduction:
1.  simplest & most primitive members of animal kingdom

   protos - first   zoon - animal

2.  range - Cambrian to Recent
3.  generally small - 1 micron to 19cm
4.  consist of a single cell
  cytoplasm; nucleus; cell wall; vacuoles
  motion by cilia, flagellum, or pseudopodia
5.  water, waste, and 02 distributed by osmosis
6.  lack systems, but are completely independent organisms (organelles)
7.  mostly solitary, but a few are colonial in protoplasm matrix
8.  feed on other protozoans and microscopic plants, plus organic debris
  some parasitic
9.  reproduction asexual or sexual - alternation of generations.  Often produce
  spores (resting stages)
10. many have some kind of external protection - hard parts
          silica - Radiolarians
          chiton (tectin)
          agglutinated material
          calcium carbonate
11. inhabit fresh, brackish, salt water, moist earth, digestive & circulatory systems
major food supply of oceans
13.  important rock formers locally
  Globigerina and radiolarian oozes
  fusulinid Ls of Penn. - Perm.
  chalks of Cretaceous & Tertiary
  Salem Ls of Indiana (Endothyrid forams)
14. very important as index fossils - especially in the petroleum industry

ORDER FORAMINIFERA
1.  possess netlike pseudopodia (extensions of protoplasm)
2.  make up 2 1/2% of all know animals, fossil or living;
  88% of the forams known as fossils
3.  mostly benthonic; shallow water to 30,000'
4.  some adapted planktonic habit in the Jurassic
  (vacuolated protoplasm, spines, reduction of S. G.)
5.  density to 2,500 per cm2 on sea floor

Skeletal features       Shell termed a TEST

1. Allogromids: Tectin - proteinaceous substance resembling
  chiton (C-Hol)
2. Textularids:  Arenaceous (agglutinated) - tests of foreign
  material.  May be highly selective as to type of material
  cements calcareous, siliceous, ferruginous (C - HOL)
3.  Calcareous -
  Rotalina: Hyaline (perforate) - (Permian - Hol)
            glassy, hole-bearing (perforate), calcite and/or
            aragonite arranged in one or more layers
      Miliolina: Porcellanous (imperforate) - (Miss - Hol)
            chalky, lack holes, tiny calcite needles, calcite xls
            parallel to outer surface layer, smooth, shiny, milky
            white surface
     Fusulinina: Microgranular  - (Ord - Tr)
            randomly arranged, equidimensional grains of calcite
            sugary appearance.   See P. 82 of Boardman/Cheetham/Rowell

Location and shape of aperture   Shape of test
figure 2.3 (Moore. et al.)   figure 2.5 (Easton)
figure 2.6 (Easton)
Ornamentation
Perforate
Imperforate
smooth surface - ornamentation
costae - ribs or ridges
granules or nodes
spines - usually planktonic
flanges
pits
reticulate - network of fine lines

Test size
micro - 0.001mm
small - few chambers (0.4mm)
larger - many chambers (2mm and larger)

Dimorphism - 2 body forms in the same species
megalospheric - asexual, few chambers, large proloculus; small test
microspheric - sexual, many chambers, small proloculus; large test
alteration of generations

Key to Superfamilies
I.  Test arenaceous  (Cambrian to Recent)
     A.  Non-chambered . . . . . . . . . . . . . . . . . . . . . . . . Astrorhizicae

     B.  Chambered . . . . . . . . . . . . . . . . . . . . . . . . . . . . Lituolicae

    II.  Test Calcareous
     A.  Mostly imperforate, non hyaline
   1.  discoid to fusiform, granular . . . .  Endothyricae
   2.  Milioline, porcelanous . . . . . . . . . Miliolicae
     B.  Mostly perforate, hyaline

  1. chambers tend to lie in linear or planispiral series,
                    particularly in early portions . . . Lagenicae
   2.  chambers tend to lie in complex whorls
    a.  tests mostly helicoid around
     a main long axis . . . . Buliminicae
    b.  tests mostly rotalioid . . . . Rotaliicae

Astrorhizicae
Arenaceous tubes which are not divided into chambers.
Recovered from insoluble residues from Ls which has been dissolved in dilute acid.

Lituolicae
Chambered arenaceous forams
Differentiated according to chamber distribution
Sil - Hol, very common in K

Endothyricae
Catch-all for granular, calcareous, planispiral forms
A few arenaceous forms
Fusulinids - larger forams, dominated the late Paleozoic
Extremely important as index fossils
Dev.(?) Miss. - Perm.
primitive forms discoidal with rounded chambers, simple walls
typified by Endothyra from the Salem Ls, Indiana
fusiform group evolved from the Endothyrids during Chester (Miss.) to give fusulinids.

Axial and equatorial sections required for I.D. wall structure of fusulinids

     Profusulinella

     Fusulina

     Schwagerina

Evolutionary trends of the Fusulinids
rapid changes in Penn. & Perm.
change in shape - discoidal to spherical, spindle-shaped, or subcylindrical
increase in size - fraction of an inch in Miss, up to 60mm in Permian
complication of shell-well structure - development of thicker primary walls and fewer wall
   layers fluting of septa
development of special structures (chromata, septula, axial fillings)
variety of internal structures

1.  Resorbed CaC03
  tunnels
  septal pores
  foramen
2.  Redeposited CaC03
  chomata - thickened zones adjacent to tunnels
  axial filling - solid central region - maximum diameter depositional sites
    for resorbed CaC03
  adjustment of S.G. with continued growth

Importance of Fusulinids
1.  Restricted geologic range - Miss. to Perm.
2.  Abundant & well preserved - fusulinid Ls
3.  Widely spread geographically - all continents except Australia & Antarctica
4.  Evolution can be traced - thin section
   80 million year life span
   72 genera, >1000 species

Miliolicae
calcareous, (porcelaneous), imperforate most coiled
resemble seeds surrounded by husks
Triassic to Holocene; abundant after mid-Cretaceous
Megalospheric & Microshperic tests; show alternation of generations
developmental series starts with:
  quinqueloculine shape, passes through a
  trioculine stage, and terminates in a
  biloculine stage
tests resemble long-extinct fusulinids

Lagenicae
calcareous, hyaline, perforate
basic shape flask-like; also uniserial & spiral
Cambrian to Recent, insignificant until Jr., when they were very common through the Cretaceous.  Still important.